Pando (Quaking Aspen)
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Pando (
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
for "I spread") is a clonal colony of an individual male
quaking aspen ''Populus tremuloides'' is a deciduous tree native to cooler areas of North America, one of several species referred to by the common name aspen. It is commonly called quaking aspen, trembling aspen, American aspen, mountain or golden aspen, tr ...
(''Populus tremuloides'') determined to be a single living organism by identical
genetic marker A genetic marker is a gene or DNA sequence with a known location on a chromosome that can be used to identify individuals or species. It can be described as a variation (which may arise due to mutation or alteration in the genomic loci) that can be ...
s and assumed to have one massive underground
root system In mathematics, a root system is a configuration of vectors in a Euclidean space satisfying certain geometrical properties. The concept is fundamental in the theory of Lie groups and Lie algebras, especially the classification and representati ...
. The plant is located in the Fremont River Ranger District of the
Fishlake National Forest Fishlake National Forest is a U.S. National Forest located in south central Utah. The namesake for the forest is Fish Lake, the largest freshwater mountain lake in the state. Wildlife Animals that inhabit this forest are elk, shrews, deer, b ...
at the western edge of the
Colorado Plateau The Colorado Plateau, also known as the Colorado Plateau Province, is a physiographic and desert region of the Intermontane Plateaus, roughly centered on the Four Corners region of the southwestern United States. This province covers an area of ...
in south-central
Utah Utah ( , ) is a state in the Mountain West subregion of the Western United States. Utah is a landlocked U.S. state bordered to its east by Colorado, to its northeast by Wyoming, to its north by Idaho, to its south by Arizona, and to it ...
, United States, around southwest of Fish Lake. Pando occupies and is estimated to weigh collectively , making it the heaviest known organism. The root system of Pando is estimated to be up to several thousand years old, placing Pando among the oldest known living organisms. Pando's long-term survival is uncertain due to a combination of factors including
drought A drought is defined as drier than normal conditions.Douville, H., K. Raghavan, J. Renwick, R.P. Allan, P.A. Arias, M. Barlow, R. Cerezo-Mota, A. Cherchi, T.Y. Gan, J. Gergis, D.  Jiang, A.  Khan, W.  Pokam Mba, D.  Rosenfeld, J. Tierney, an ...
, human development,
grazing In agriculture, grazing is a method of animal husbandry whereby domestic livestock are allowed outdoors to roam around and consume wild vegetations in order to convert the otherwise indigestible (by human gut) cellulose within grass and other ...
, and
fire suppression Wildfire suppression is a range of firefighting tactics used to suppress wildfires. Firefighting efforts in wild land areas require different techniques, equipment, and training from the more familiar structure fire fighting found in populated a ...
. The Western Aspen Alliance is studying the tree in an effort to save it in collaboration with the
United States Forest Service The United States Forest Service (USFS) is an agency of the U.S. Department of Agriculture that administers the nation's 154 national forests and 20 national grasslands. The Forest Service manages of land. Major divisions of the agency inc ...
. In areas of Pando lacking adequate protective fencing, grazing animals have prevented Pando from developing enough young stems to fully replace existing older stems as they die. This decline in recruitment of young stems began in the 1980s and has been attributed primarily to
mule deer The mule deer (''Odocoileus hemionus'') is a deer indigenous to western North America; it is named for its ears, which are large like those of the mule. Two subspecies of mule deer are grouped into the black-tailed deer. Unlike the related whit ...
(''Odocoileus hemionus''), though Pando also undergoes restricted grazing by
domestic cattle Cattle (''Bos taurus'') are large, domesticated, cloven-hooved, herbivores. They are a prominent modern member of the subfamily Bovinae and the most widespread species of the genus '' Bos''. Adult females are referred to as cows and adult m ...
(''Bos taurus'') and is potentially browsed by
elk The elk (''Cervus canadensis''), also known as the wapiti, is one of the largest species within the deer family, Cervidae, and one of the largest terrestrial mammals in its native range of North America and Central and East Asia. The common ...
(''Cervus canadensis'').


History

Pando was identified in 1976 by Jerry Kemperman and Burton Barnes. Michael Grant, Jeffrey Mitton, and Yan Linhart of the
University of Colorado at Boulder The University of Colorado Boulder (CU Boulder, CU, or Colorado) is a public research university in Boulder, Colorado. Founded in 1876, five months before Colorado became a state, it is the flagship university of the University of Colorado sys ...
re-examined the clone in 1992, naming it Pando and claiming it to be the world's largest organism by weight. Both teams of researchers described Pando as a single asexual reproduction organism based on its morphological characteristics. Genetic sampling and analysis in 2008 by Jennifer DeWoody, Carol Rowe, Valerie Hipkins, and Karen Mock of
Utah State University Utah State University (USU or Utah State) is a public land-grant research university in Logan, Utah. It is accredited by the Northwest Commission on Colleges and Universities. With nearly 20,000 students living on or near campus, USU is Utah's ...
and the
University of Southampton , mottoeng = The Heights Yield to Endeavour , type = Public research university , established = 1862 – Hartley Institution1902 – Hartley University College1913 – Southampton University Coll ...
confirmed the earlier analyses and increased the clone's estimated size from 43.3 to 43.6 ha. Paul Rogers and Darren McAvoy, also with Utah State University, completed the first comprehensive assessment of Pando's status in 2018 and stressed the importance of reducing herbivory by mule deer to conserving Pando for the future. Rogers and Jan Šebesta surveyed other vegetation within Pando besides aspen in 2019, finding additional support for their 2018 conclusion interactions between browsing and past and ongoing management have had adverse effects on Pando's long-term resilience to change. While Pando is the largest known aspen clone and its age has received considerable attention, other large and old clones exist. According to an
Organisation for Economic Co-operation and Development The Organisation for Economic Co-operation and Development (OECD; french: Organisation de coopération et de développement économiques, ''OCDE'') is an intergovernmental organization, intergovernmental organisation with 38 member countries ...
(OECD) report in 2000: In 2006 the
United States Postal Service The United States Postal Service (USPS), also known as the Post Office, U.S. Mail, or Postal Service, is an independent agency of the executive branch of the United States federal government responsible for providing postal service in the U ...
published a stamp in commemoration of the aspen, calling it one of the forty "Wonders of America".


Size and age

The clonal colony encompasses , weighs nearly , and has over 40,000 stems (trunks), which die individually and are replaced by new stems growing from its roots. The root system is estimated to be several thousand years old with habitat modeling suggesting a maximum age of 14,000 years. Individual aspen stems typically do not live beyond 100–130 years and mature areas within Pando are approaching this limit. Mitton and Grant summarize the development of stems in aspen clones:


Range of age estimates

Due to the progressive replacement of stems and roots, the overall age of an aspen clone cannot be determined from
tree ring Dendrochronology (or tree-ring dating) is the scientific method of dating tree rings (also called growth rings) to the exact year they were formed. As well as dating them, this can give data for dendroclimatology, the study of climate and atmos ...
s. In Pando's case, ages up to 1 million years have therefore been suggested. An age of 80,000 years is often given for Pando, but this claim derives from a
National Park Service The National Park Service (NPS) is an agency of the United States federal government within the U.S. Department of the Interior that manages all national parks, most national monuments, and other natural, historical, and recreational propertie ...
web page that does not provide a source for its number and is inconsistent with the Forest Service's post ice-age estimate. Glaciers repeatedly formed on the Fish Lake Plateau over the past several hundred thousand years and the Fish Lake valley occupied by Pando was partially filled by ice as recently as the
last glacial maximum The Last Glacial Maximum (LGM), also referred to as the Late Glacial Maximum, was the most recent time during the Last Glacial Period that ice sheets were at their greatest extent. Ice sheets covered much of Northern North America, Northern Eur ...
. Ages greater than approximately 16,000 years therefore require Pando to have survived at least the Pinedale glaciation, something that appears unlikely under current genetic estimates of Pando's age and modeling of variation in Pando's local climate. Estimates of Pando's age have also been influenced by changes in the understanding of establishment of aspen clones in western North America. Earlier sources argued germination and successful establishment of aspen on new sites was rare in the last 10,000 years and, therefore, Pando's root system was likely over 10,000 years old. More recent observations, however, have shown seedling establishment of new aspen clones is a regular occurrence and can be abundant on sites exposed by
wildfire A wildfire, forest fire, bushfire, wildland fire or rural fire is an unplanned, uncontrolled and unpredictable fire in an area of Combustibility and flammability, combustible vegetation. Depending on the type of vegetation present, a wildfire ...
. These findings are summarized in the U.S. Forest Service's ''Fire Effects Information System'':


See also

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Basal shoot Basal shoots, root sprouts, adventitious shoots, and suckers are words for various kinds of shoots that grow from adventitious buds on the base of a tree or shrub, or from adventitious buds on its roots. Shoots that grow from buds on the base of ...
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List of oldest trees This is a list of the oldest-known trees, as reported in reliable sources. Definitions of what constitutes an individual tree vary. In addition, tree ages are derived from a variety of sources, including documented "tree-ring" (dendrochronologica ...
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List of individual trees The following is a list of notable trees. Trees listed here are regarded as important or specific by their historical, national, locational, natural or mythological context. The list includes actual trees located throughout the world, as well as ...
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Rhizome In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow hori ...
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Vegetative reproduction Vegetative reproduction (also known as vegetative propagation, vegetative multiplication or cloning) is any form of asexual reproduction occurring in plants in which a new plant grows from a fragment or cutting of the parent plant or spec ...
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Largest organisms The largest organisms now found on Earth can be determined according to various aspects of an organism's size, such as: mass, volume, area, length, height, or even genome size. Some organisms group together to form a superorganism (such as ants ...


References


Additional references from OECD quote in history section

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External links

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Pando
at
Atlas Obscura ''Atlas Obscura'' is an American-based online magazine and travel company. It was founded in 2009 by author Joshua Foer and documentary filmmaker/author Dylan Thuras. It catalogs unusual and obscure travel destinations via user-generated content ...
{{Portal bar, Utah Populus Individual trees in Utah Fishlake National Forest Oldest trees